It's Christmas, and Angelina Mouseling (Finty Williams) is auditioning for the lead role in the musical extravaganza, Cinderella Mouse and the Ugly Whiskers. She is devastated when Penelope Pinkpaws (Jonell Elliott) is chosen to play Cinderella and she is cast as the Wicked Stepmouse. Taking Miss Lilly's (Dame Judi Dench's) advice to "think horrid" much too literally, Angelina storms off in a fit of a temper. Then a toboggan race goes wrong and it seems that the production will have to be cancelled. Realizing how selfish she has been, Angelina determines to sort things out, because as every aspiring star knows, "The Show Must Go On".